I'm trying Seamonkey ... it has a much smaller RAM footprint than running Firefox and Thunderbird together. Seems pretty clean (uses the same open source but if you still run an email client it saves a lot of space by sharing libraries I'm assuming).
Only "problem" I've encountered so far is that it has some different defaults than FF and TB, so you have to hunt those down in the options. It has the necessary pile of addons (e.g. flashgot, adblocker, etc) but isn't quite as "pretty" as firefox can be made to be.
Seems to be a good choice for older machines with limited RAM (1GB or less) and they're an active free opensource project.
edit: so far looking wonderful... will post if I encounter any issues.
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