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Kiroumaru and Yakomaru DO have some pretty strong similarities, shown from this reveal. They're both willing to be dishonest and show loyalty to the humans while plotting in secret. The big difference between them is that Kiroumaru seems to value a peaceful status quo and Yakomaru is looking to carve out an empire of his own. Even then, we're giving Kiroumaru the benefit of the doubt in saying this - we don't know what his course of action would be if he had actually found WMDs when visiting.
The main differences lie in their personalities, I think. Yakomaru doesn't care for giving an air of chivalry and will do whatever it takes. Kiromaru does value these traits, but as a leader his loyalty was to his people above his loyalty to humanity, so he felt it was more 'noble,' if you will, to plot in secret. In both situations, I think humanity was asking for trouble with the way they lord over the rats. Yakomaru seems to be a Julius Ceasar of sorts; taking advantage of the hostile climate to propel himself into a position of prominence, but there's no denying that if you were in either colony, the humans must have seemed to be a huge threat to your well-being.
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