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The fact that Nayuki appears first, is the best developed female character, arguably has the most involving story, and has the most screen time all seemed to imply that she was the actual main character in the game.
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Now if that were true all the time, I wouldn't have had problems with Shuffle either. XD
But seriously though, I see Ayu more than Nayuki in promotional stuff from Key. In one calendar (was it?), it was Misuzu from Air, Nagisa from Clannad, and you guessed it, Ayu from Kanon in the front. I also vaguely remember an older picture from KEY that included Misuzu and Ayu, but had Mizuka from ONE also. So I always had thought that Ayu was the lead of Kanon.