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Basically:
Let's call the usual twins we know with their appropriate name, while their "place" will be "heir" and "little sister".
Originally, Shion (the long haired we know all the time) was the true "heir" of the Sonozaki, as being the elder of the twins, the first born.
However, both Shion and Mion disapproved how the "little sister" slot ("shion") is in a unfair situation. So both twins just swap all the time so both can enjoy the "Mion" (so "heir") position.
Unfortunately, one day, Shion was "shion" (so the little sister) and was oddly invited to a amusement park or something. Mion (who was supposed to the shion, the true little sister), received the tatoo, the proof of the heir of the sonozaki.
Because of this, Mion and Shion were forced to play officially their opposite original positions. Mion became "Mion" while she was supposed to be "Shion".
This is the reason why Mion was always guilty and feelign sorry for Shion, because not only she wasn't exactly well treated, but she kinda "stole" Shion's happiness. That is the reason that, when Shion became nuts, they "naturally" reverted to their true positions, and Shion literally became "Mion" (the heir).
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