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I've been thinking:
Horo's non-sexualized nudity is a depiction of her divinity.
Before Christianity took over Europe, most of the classical Cultures of Europe and the Mediterranean world depicted divine figures in the nude. Greek and Roman gods were depicted in the nude, 'divine' and 'semi-divine' humans such as 'divine' kings and emperors were depicted in the nude in art and sculpture when their mortal counterparts were clothed.
Horo's nudity is a representation of her divine or semi-divine nature and because of that if anyone accuses fans of Spice & Wolf of being perverts just looking at Horo, we can explain that there are very good symbolic reasons why Horo is frequently depicted in the nude in a non-ecchi way...
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