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Though to be fair, I also thought Kanon was a girl pretty much until he died in Episode 4 of the anime, and he has the only "androgynous" sounding voice, to me. I don't think I'd believe Lion's voice as a boy, and Zepar's voice sounds too ... exaggerated? Though they're all voiced by women, so, /shrug
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Hah, Same here I thought Kanon was a girl when I first watched anime, I even thought 'Heh What a cool girl' and then I facepalmed myself